Abu Layla, who commanded a brigade inside the predominantly-Kurdish Syrian Democratic Forces, was hit by IS sniper fire on the outskirts of Manbij, an Islamic State group stronghold that controls the supply route between the Turkish border and Raqqa, the Kurdish website Rudaw said. He was evacuated by US forces to a hospital in the Iraqi Kurdish city of Sulaymaniyah, where he died.
The commander fought against IS militants in Kobani in early 2015, according to the Britain-based Syrian Observatory for Human Rights, a monitoring group. Those battles, the first major setback to the IS advance in northern Syria, were seen as instrumental to securing US support for Kurdish forces in the country's multi-layered conflict.
Pro-government forces reached within 40 kilometers (25 miles) of the Tabqa Air Base, to the west of Raqqa, according to the Observatory.
IS militants captured the base from the government in 2014, killing scores of captured soldiers. The media arm of Lebanon's Hezbollah group, which is fighting alongside government forces, said the Syrian army took a small village 50 kilometers (31 miles) to the southwest of Tabqa air base.
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