Military prosecutors are investigating allegations of secret illegal contacts between Polish and Russian intelligence officials during the period when Tusk was Poland's prime minister.
He told reporters as he left the prosecutors' office in Warsaw that he cannot give details about his closed-door testimony, which is secret, but that he was treated hospitably to "tea, coffee and water."
"The whole case has a highly political character," Tusk said.
Many see his questioning as part of a larger attempt by Poland's nationalist government to discredit a political foe by linking him to scandals.
With backing for Tusk still strong in the country, he could prove to be a serious rival to the ruling party in the 2019 parliamentary election and in the 2020 presidential election.
Tusk has also been accused by Defense Minister Antoni Macierewicz of treason in another matter, the handling of the aftermath of the 2010 plane crash in Smolensk, Russia, that killed Polish President Lech Kaczynski.
Emotions at the train station ran high. Tusk's supporters carried EU and national flags and chanted, "Donald, we are with you!" Opponents hurled accusations of crimes and of Tusk having hurt Poland's interests. One detractor raised a large mock-up photo depicting him in striped prison garb.
In the current case, prosecutors have not revealed details of the alleged illegal contacts, but they are investigating whether the heads of Poland's intelligence neglected their obligation to seek Tusk's approval for cooperating with foreign intelligence agencies.
"I have no reservations as to the work of the (special) services," Tusk said before his meeting with prosecutors. Poland's current ruling conservative party, Law and Justice, is led by the late president's twin brother, Jaroslaw Kaczynski, a nationalist politician who is a long-term political rival of Tusk's.
Kaczynski and others accuse Tusk of failing to oversee proper security for the presidential flight. They also fault Tusk for letting the Russians carry out the main investigation and for failing to get the wreckage back.
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