Pyotr Levashov, a 37-year-old from St Petersburg, was arrested in April while vacationing with his family in Barcelona.
US authorities had asked for him to be detained on charges of fraud and unauthorized interception of electronic communications. He was scheduled to be arraigned late Friday in a federal courthouse in Bridgeport, Connecticut, where a grand jury indicted him last year.
A statement from Spain's National Police said officers handed Levashov over to US marshals yesterday. The extradition was approved in October by Spain's National Court, which rejected a counter-extradition request from Russia.
Authorities in the US say they have linked Levashov to a series of powerful botnets, or networks of hijacked computers, that were capable of pumping out billions of spam emails.
An indictment unsealed last year said he commanded the sprawling Kelihos botnet, which at times allegedly involved more than 100,000 compromised computers that sent phony emails advertising counterfeit drugs, harvested users' logins and installed malware that intercepted bank account passwords.
On a typical day, the network would generate and distribute more than 2,500 spam emails, according to the indictment.
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