"If the Russian army kills the people of Syria, then kill their people. And if they kill our soldiers, then kill their soldiers. An eye for an eye," Abu Mohamed al-Jolani, the head of Al-Nusra Front, said in an audio recording released late yesterday.
He pledged that Moscow's air war in Syria, which began on September 30, would have dire consequences for Russia.
"The war in Syria will make the Russians forget the horrors that they found in Afghanistan," Jolani said, adding: "They will be shattered, with God's permission, on Syria's doorstep."
State group, a jihadist rival of Al-Nusra, as well as other extremist groups.
Moscow's strikes have concentrated on areas in northwestern Syria where Al-Nusra has a powerful presence, mostly in the province of Idlib.
A US-led air coalition fighting IS in Syria has also targeted Al-Nusra on several occasions.
Jolani called on armed opposition groups to set aside their differences until both air wars had been defeated.
"Delay the disputes until the demise and smashing of the Western Crusader and Russian campaign on Syrian land," he said.
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