Authorities began destroying living quarters at Larung Gar Institute in the southwestern province of Sichuan last week, according to a statement by the International Campaign for Tibet (ICT).
In June, local government officials ordered the estimated 10,000 Buddhist monks and nuns living at the centre, also known as Serthar, be cut to no more than 5,000, another overseas organisation, Free Tibet, said in an email.
"These demolitions are part of a set of policy measures implemented by the Chinese government that severely restrict the religious freedom of Tibetan Buddhists," wrote the ICT's president Matteo Mecacci.
Serthar, known as Seda in Chinese, is in an ethnically Tibetan area more than 4,000 metres above sea level and hundreds of kilometres from the nearest city.
The institute was founded in 1980 in an uninhabited valley and has since grown to become one of the world's most important centres of Tibetan Buddhism, with monks, nuns and students living in small wooden homes sprawling over the hillsides surrounding the complex.
China accuses the Nobel peace laureate of fomenting unrest in the region, but overseas rights groups say incidents are driven by unhappiness at Beijing's rule among Tibetans.
Pictures circulated on social media showed workmen and "mechanical diggers" destroying homes at the institute, the ICT statement said.
A local propaganda official surnamed Ni told AFP that the work was intended to "prevent fires" by giving firefighters access to the crowded area and had "received the monks' active cooperation".
The Buddhist academy's survival was threatened in 2001 when armed police forced hundreds of nuns and monks to leave the site, destroying more than 1,000 homes to prevent them returning.
At the time police demanded that nuns sign documents denouncing the Dalai Lama and pledging not to return, according to overseas campaign groups, and Khenpo Jigme Phuntsok was detained for a year after the incident.
China's official Xinhua news agency said the incident, in which one person was killed, was triggered when "rioters" attacked police.
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