He will be seen as the father of villain The Penguin, reported Deadline.
Robin Lord Taylor, who plays the penguin, shared the news at the New York Comic-Con.
Reubens, 63, also played the father of The Penguin 23 years ago in Tim Burton's "Batman Returns". That Penguin was played by Danny DeVito.
The series is based on characters appearing in and published by DC Comics in their Batman franchise, primarily those of James Gordon and Bruce Wayne.
Ben McKenzie stars as young Gordon. The cast also includes Cory Michael Smith and Erin Richards plus new members Michael Chiklis, James Frain and Jessica Lucas.
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