The seven-match tour has been spiced up by comments from England captain Alastair Cook that his team will try to exact revenge for the Test and one-day defeats at home.
Cook's status as captain came under heavy pressure after the 1-0 Test defeat, and tempers frayed in the ODIs when off-spinner Sachithra Senanayake controversially ran out Jos Buttler before delivering a ball.
"There is a World Cup at the end of it, but we are here to try and win this series," Cook said on arrival. "I think we owe you (Sri Lanka) a little bit of revenge."
But Sri Lanka insisted Buttler was earlier warned for backing up, and senior pro Mahela Jayawardene lashed out at Cook for questioning Senanayake's run-out.
Meanwhile Sri Lanka are smarting from this month's 5-0 thrashing in India in a series that was hastily arranged after the West Indies abandoned their Indian tour over a pay dispute.
Sri Lankan captain Angelo Mathews insisted the matches will be used to recover from the morale-sapping loss and find the right combination for the World Cup starting in February.
"It's very important that we sort out our bowlers and also the batting order. By the end of the series we should have a very clear idea of whom we want to take to the World Cup."
Both teams are grouped together in the preliminary stage of the World Cup and will feature on the opening day on February 14, with Sri Lanka playing New Zealand and England facing Australia.
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