Abu Firas al-Suri, whose real name was Radwan Nammous, fought against Soviet forces in Afghanistan where he met Al-Qaeda leader Osama bin Laden and the founding father of global jihad, Abdullah Azzam, before returning to Syria in 2011.
Suri was meeting with other leading Islamist fighters in an Al-Nusra stronghold in Kafar Jales in northwestern Syria when the raids struck yesterday.
He "was an old time Al-Qaeda member ... He was brought in from Yemen as an ideological counterweight" for rival jihadist group IS, said Pieter Van Ostaeyen, a historian and monitor of jihadist groups.
According to the Syrian Observatory for Human Rights, Suri, his son and at least 20 jihadists of Al-Nusra and Jund al-Aqsa and other fighters from Uzbekistan were killed in strikes on positions in Idlib province.
Seven were high-ranking jihadists, the Britain-based Observatory said, adding that the Syrian air force had likely carried out the strikes.
A temporary ceasefire between government forces and rebels has largely held since February 27, but it does not cover Al-Nusra and IS.
The break has, in fact, allowed Russia and the US-led coalition that has been bombing IS in Syria to concentrate on their fight against the jihadists.
But on Friday, the Al-Qaeda affiliate and allied rebel groups pushed regime loyalists out of Al-Eis, a strategic town in the northern province of Aleppo, killing 12 members of the Lebanese Shiite Hezbollah movement.
"It was Al-Nusra's biggest operation since the ceasefire began," Observatory chief Rami Abdel Rahman said.
Suri's killing may even be a warning by the regime to Al-Nusra against staging any more offensives in future.
Al-Nusra's rival IS has also lost a string of high-ranking members in recent weeks, mainly to strikes by the US-led coalition that launched an aerial campaign against the jihadists in Iraq and Syria in 2014.
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