Sudanese vets scrambled to save four surviving captive lions of five that were left sick and malnourished in a Khartoum park during months of political and economic turmoil.
The fifth, a lioness, died on Monday of dehydration and lack of food despite receiving intravenous fluids for several days following the launch of an online campaign for veterinary help.
The animals had lost as much as two-thirds of their body weight as a result of going unfed for weeks as Sudan's chronic cash shortage hit donations.
"It's so upsetting. These lions have been suffering for so long," said Osman Salih, a software engineer who launched the online campaign under the hashtag #Sudananimalrescue.
"We were so late in rescuing them and were unable to save the one that died."
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